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A large proportion of the land surface of Africa bears vegetation in which grass is an essential feature. This abundance of grass has made possible the continent’s enormous and varied populations of herbivorous mammals, both wild and domesticated. The tall and fibrous invasive grasses in forest environments and in large tracts of wooded grasslands are seldom very palatable to livestock, but, in those parts of the continent where good forage grows naturally or has been introduced, livestock raising is of great economic importance.

The Albida acacia tree of the “farmed parkland” areas of western Africa is of special economic importance. Unlike almost all other dry woodland trees, whose leaf shedding normally occurs at the onset of the dry season, the Albida appears to have a period of partial dormancy during the rainy season and springs to life only at the beginning of the dry season. At such periods its foliage is abundant and—being a palatable leguminous species—is much prized as browse for sheep, goats, and cattle. The smaller leafy branches are frequently fed to stock. The tree flowers and produces fruits, which are harvested, dried, and fed as a protein concentrate to stock at the height of the dry season. Animal resources

Water buffalo, oxen, horses, mules, donkeys, and camels are used primarily as draft, pack, or riding animals in Africa, and they also provide milk, meat, hides, or skins. Because of its intractability and wild nature, however, the African elephant—unlike the Asian elephant—is not used for draft or haulage purposes.

The water buffalo is an offshoot of the Asiatic buffalo (Bubalus bubalis); it arrived in Africa in relatively recent times and is now found almost exclusively in Egypt. The domesticated African water buffalo is used to cultivate irrigated land (mainly in the Nile delta) and to provide milk and meat.

Oxen are widely used in Africa for agricultural purposes, especially for plowing and cultivation; they are also trained to thresh grain, pump water, and act as pack animals. Bullock (castrated oxen) plowing is well developed in the countries of North Africa, in Ethiopia and Sudan, and farther west in Chad, in northern Nigeria, and in the savanna climatic zone of western Africa. Plowing and cultivation by oxen is also well developed in areas of eastern and Southern Africa that are free of the deadly tsetse fly. Females used for work may also be milked. Work oxen are often used for meat and to provide hides.

Horses and ponies are principally found in Morocco, Algeria, Tunisia, Chad, Ethiopia, Mali, Niger, Nigeria, Senegal, Burkina Faso, Egypt, South Africa, and Lesotho. Horses are used as riding or pack animals and in a number of areas are bred with donkeys to produce mules. Few are kept in areas where tsetse flies are present. Five main types of horses inhabit Africa: the Darfur pony, the Dongola horse, the Ethiopian-Galla horse, the Somalia pony, and the South African horse (including the Basuto pony). In North Africa, types also have evolved as a result of selection and crossing with exotic Arab, Barb (Barbary), and Thoroughbred horses. Arab and Thoroughbred influence may also be noted in Southern Africa.

The distribution of the ass roughly corresponds to that of the horse, except that it also extends into the livestock areas of eastern and central Africa. Mules are found in Algeria, Ethiopia, Morocco, Somalia, South Africa, and Tunisia, where they provide farm draft power and are used as pack animals and for riding. The ability of mules to perform work in hot, dry climates is superior to that of most other farm animals.

The Arabian camel, or dromedary, is widely dispersed in the drier regions of northern and eastern Africa. Although used principally as a pack animal, it also is used for land cultivation, water pumping, and human transportation. The camel is essentially a bush browser and, if reasonably well fed and watered, may produce about 11 to 13 pounds (5 to 6 kilograms) of milk daily, in addition to that fed to the calf. The milk is prized by the camel herders and their families. Camel meat and camel hides find a ready market among Muslim communities.

Cattle provide hides, and sheep, goats, and pigs provide skins. Skins of the Maradi, Sokoto, and Kano red goats from Niger and Nigeria are greatly prized by the Morocco leather trade. In the areas north and south of the tropical zone, African sheep are covered with wool, but in the tropics they are hairy. In elevated areas, such as Ethiopia, where temperatures are modified by altitude, some sheep may be partially wooled, at least on the back and buttocks. The wooled sheep of North Africa are largely of the woolly Barbary type, which was originally introduced to Africa from the Middle East.

The great herds of wild African herbivores include the principal game animals. African antelope have been important throughout human history as sources of meat and such by-products as hides and bone, and they, along with other large mammals, became prized by trophy hunters. For centuries African elephants were sought for the ivory in their tusks, but the severe reduction of their numbers by the late 20th century led to a total ban on hunting them in most African countries.

The most economically useful fishes found in African waters include many freshwater species. Important among the marine fishes are flounder, halibut, sole, redfish, bass, conger, jack, mullet, herring, sardine, and anchovy. Crustaceans are important for local consumption and for export, as are oysters (for pearls), trochus shells, corals, and sponges. The most economically important aquatic mammal is the Southern, or Cape, fur seal. Agriculture

Agriculture is by far the single most important economic activity in Africa. It provides employment for about two-thirds of the continent’s working population and for each country contributes an average of 30 to 60 percent of gross domestic product and about 30 percent of the value of exports. Nonetheless, arable land and land under permanent crops occupy only about 6 percent of Africa’s total land area.

Except for countries with sizable populations of European descent—such as South Africa, Zimbabwe, and Kenya—agriculture has been largely confined to subsistence farming and has been considerably dependent on the inefficient system of shifting cultivation, in which land is temporarily cultivated with simple implements until its fertility decreases and then abandoned for a time to allow the soil to regenerate. In addition, over most of Africa arable land generally has been allocated through a complex system of communal tenure and ownership rather than through individually acquired title, and peasant farmers have had rights to use relatively small and scattered holdings. This system of land ownership has tended to keep the intensity of agricultural production low and has inhibited the rate at which capital has been mobilized for modernizing production. A number of countries have made efforts to raise productive levels by selecting better varieties of seeds and planting materials, using tractors and other mechanized equipment, or increasing the use of mineral fertilizers and insecticides. Such measures, however, have been relatively limited, and they have raised concerns about their part in accelerating soil erosion and desertification. In areas of cash crop production, land has become private rather than community property, and cultivation is intensive.

The persistence of relatively low-productivity agricultural systems over large parts of the continent also stems from a lack of integration between crop production and animal husbandry. Traditionally, sedentary cultivators like the Hausa in Nigeria and the Kikuyu in Kenya live apart from their nomadic herdsmen neighbours (the Fulani and Maasai, respectively), with the result that over large areas of the continent farmers do not have access to animals for draft power or to manure for fertilizer. The incidence of such insect pests as the tsetse fly also discourages mixed farming in many areas.