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But for now, Noth was in no position to challenge the Emperor or any of the stronger males in the loose pecking order above him. And he could see that the supply of podocarp fruit was dwindling rapidly.

With a frustrated hoot he hurried over the forest floor and scampered briskly into a tree. The branches, slippery with residual frost, dew, and lichen, were all but bare of leaves and fruit. But it might still be possible to find caches of nuts or seeds, stashed away by providential forest creatures.

He came to a hollow in an aging tree trunk. In its dank, rotting interior, he saw the gleam of nutshells. He reached in with his small, agile hands and hauled out one of the nuts. The shell was round, seamless, complete. When he rattled it he could hear the kernel inside, and saliva spurted into his mouth. But when he bit into the shell his teeth slid over the smooth, hard surface. Irritated, he tried again.

There was an almighty hissing. He hooted, dropped the nut, and scuttled to a higher branch.

A creature the size of a large domestic cat came scrambling clumsily toward the nut cache. It raised its head to Noth and hissed again, showing a pink mouth with powerful upper and lower incisors. Satisfied that it had driven off the raider, it dug out one of its stored nuts and, with a clench of its powerful jaws, cracked the shell. Soon it was nibbling purposefully, widening the hole it had made. At last it reached the nut’s kernel — Noth, tucked behind the tree trunk, was almost overwhelmed by the sudden sweet aroma — and fed noisily.

This ailuravus looked something like a rudimentary squirrel, with a mouselike face. It had a long bushy tail, the purpose of which was to slow its fall, parachute-like, every time it tumbled out of a tree, as it frequently did. Although it was clumsier than the notharctus as it moved about in the trees, lacking a primate’s grasping hands and feet, it was more than big enough to have fought off Noth.

The ailuravus was one of the first rodents. That vast, enduring family had emerged a few million years earlier in Asia, and had since migrated around the world. This small encounter was a skirmish at the start of an epochal conflict for resources between the primates and the rodents.

And the rodents were already winning.

They were beating primates to the food, for one thing. Noth would have needed a nutcracker to eat hazelnuts or brazil nuts, and a millstone to process grains like wheat and barley. But the rodents, with their ferocious, ever-growing incisors, could break through the toughest nut and grain seed coats. Soon they would begin to consume the fruits of the best trees before they were even ripe.

Not only that, the rodents outbred the primates by a large factor. This ailu could produce several litters within a single year. Many of its young would fall to starvation, competition with their siblings, or predation by birds and carnivores. But it was enough that some should survive to continue the line, and for the ailu each of its young represented little investment — unlike the notharctus who bred just once a year and for whom the loss of a single pup was a significant disaster. And the rodents’ vast litters incidentally offered up much raw material to the blind sculptors of natural selection; their evolutionary rate was ferocious.

Even though primates like Noth were much smarter than rodents like the ailu, his kind could not compete.

It wasn’t just the plesiadapids that were becoming rare in North America. It was no coincidence that Noth’s kind had been pushed up into this marginal polar forest. In the future Noth’s line would migrate further, passing over the roof of the world to Europe and thence to Asia and Africa, adapting and reshaping as they went. But in North America the rodent victory would, within another few million years, be complete. A new ecology would arise, populated by gophers, squirrels, pack rats, marmots, field mice, and chipmunks. There would be no primates in North America: none at all, not for another fifty-one million years, not until human hunters, very distant descendants of the notharctus, came walking over the Bering Strait from Asia.

When the rodent was done feeding, Noth crept cautiously out of his hiding place. With his agile hands he sought out the scraps of kernel the ailu had dropped, and crammed them into his mouth without shame.

For a few hours a day the southern sky still grew bright. But the sun made its cycles beneath the horizon now. Almost all of the lakes were frozen over, and the trees were laden with frost, some of it gleaming in thick lacy shards where mist had frozen out on spiderwebs. The notharctus’ movements through the trees and over the silent forest floor were sluggish and dull. But it didn’t matter; the forest could offer them little more food this autumn.

There came a last clear day, when layers of red-tinged cloud stacked up against a violet southern sky, and the purple-green aurora rolled like a vast curtain over the stars.

The notharctus hurried to the ground and began to dig in places where the soil had been kept unfrozen by layers of leaves or under the roots of trees. Tonight would be the hardest frost of the winter so far, and they all knew it was time to get under cover. So the primates dug, building burrows in which Purga would have felt comfortable. It was as if the brief interval in the trees had been nothing but a dream of freedom.

In the deepest dark, Noth pushed his way through tunnels quickly worn smooth by the passage of primate bodies and over a floor littered with loose fur. At last his powerful nose guided him to Right.

Gently Noth sniffed his sister. She was already dormant, curled tightly up with her tail wrapped around her, close to the belly of Big. She had grown during their months with Biggest’s troop, but Right would always be small, always retain traces of the runt who had been bullied by her now-dead twin. Still her winter fur seemed sleek, healthy, and free of knots and dirt, and her tail was fat with the store that should sustain her through the winter.

Noth felt a kind of satisfaction. Given their dreadful start to the summer, they had both beaten the survival odds better than expected. With no pups of his own, this was still the only kin Noth had — his entire genetic future depended on Right — but for now there was nothing more he could do for her.

In darkness, immersed in the scents and subtle noises of his kind, Noth snuggled as close as he could to his sister. He shut his eyes, and was soon asleep.

Briefly he dreamed, of fragments of summer light, of long shadows, of his mother’s fall from the trees. And then, as his body shut down, his mind dissolved.

IV

The sun’s rays, almost horizontal, shone like searchlights into the forest. Above the slowly melting ponds a chill mist hovered, shining in elaborate pink-gray swirls, pointlessly beautiful. From the gaunt tree trunks immense black shadows stretched north. But the first leaves were budding on the bare branches, tiny green plates hanging almost vertically to catch the sun’s light. The leaves were already at work: The days of spring and summer were so short that these hardy vegetable servants had to gather every droplet of light they could.

It was just a glimpse, a dawn that would last no more than a few minutes. But it was the first time for several months that the disk of the sun had shown.

The forest was quiet. The great herbivorous migrants were still hundreds of kilometers to the south; it was weeks yet before they would come back, seeking their summer feeding grounds, and even the birds had yet to arrive. But Noth was already awake, already out, working.