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The males’ negotiations were more dramatic.

Whiteblood found himself facing two younger males, brothers, in fact. One had a peculiar crest of snow-white hair that stuck up around his eyes, making him look permanently surprised, and the other had a habit of using his left arm predominantly over his right, so much so that the muscles on his left side were much more heavily developed than those on the right, like those of a left-handed tennis player.

Both Crest and Left were smaller and weaker than Whiteblood, and, younger, they had not outranked him back in the forest. But now Whiteblood had lost all of his allies, and together these two might defeat him.

So, without hesitation, he launched into a display. He stood upright, shakily, hooted and shrieked, and threw handfuls of leaves. Then he turned around, spread his backside and blew shit through moist fur.

Left was immediately intimidated. He shrank back, arms folded around himself.

Crest was more defiant, and answered Whiteblood’s display with a shrieking tantrum of his own. But he was outsized by Whiteblood and, without the support of his brother, could not hope to best the older male. When Whiteblood began to cuff him about the head and neck, Crest quickly backed down, tumbling onto his back and spreading his arms and legs like an infant, showing his submission. All of this was halted only when an incautious stamp plunged Whiteblood’s leg through the foliage and into the cold water. He yelped, pulled back his leg, and sat with legs folded beneath him, subdued.

But he had done enough. The brothers approached him now, their heads bent and postures humble. A brief interval of frantic mutual grooming ensured the new hierarchy was reinforced, and the three males started to pick bits of shit out of each other’s fur.

The rough-and-ready communities of Noth had been like street gangs, held together by not much more than brute force and dominance, with each individual aware of little more than her own place in the pecking order. But by now the advantages of social living had driven primate societies to baroque intricacy, and had spurred the development of new types of mind.

Group living required a lot of social knowledge: knowing who was doing what to whom, how your own actions fit in with this, who you had to groom and when, to make your life easier. The larger the group, the greater the number of relationships you had to keep track of, and as those relationships changed constantly, you needed still more computational capacity to handle it all. By allowing their group living to develop to such extremes of complexity, primates continued to get relentlessly smarter.

Not all primates, though.

Through all this the big potbelly had sat on the comfortable branch she had found, methodically stripping it of leaves. She had no interest in the peculiar displays and hairy fiddling of the anthros.

Even among her own kind the potbelly knew little of the society of others. She ignored other females and let herself be bothered by males only when she felt the urge to mate — which, in fact, was on her now. When they were in season anthros like Patch and Roamer showed sexual swellings on their rumps. That would have been of little use to a creature who spent most of her time sitting on her backside, so on the potbelly’s chest pinkish blisters had swollen brightly in an unmistakable hourglass shape. But as there was no male potbelly around, nobody was doing anything about it.

Not that the potbelly cared much. She didn’t understand where she was and what had become of her any more than the anthros did, but it didn’t trouble her. She could see there were plenty of leaves on this fallen tree to last her through the day. She had no real idea that there could be such a thing as a tomorrow different from today, that it might not find her in an endless forest full of nutritious leaves.

Already the anthros were starting to feel hungry; their low-nutrition diet worked through their systems quickly. They broke up their grooming circles and spread out over the branches of the fallen mango. The tree had lost much of its fruit, along with most of its inhabitants, when it fell from the bank. But Crest, one of the brothers, quickly turned up a cluster of fruit that had gotten lodged in an angle of branch and trunk. He hooted to summon the others.

The new miniature society worked efficiently. Though Crest managed to grab one piece of fruit for himself, he was quickly pushed away by Whiteblood. But Whiteblood was in turn usurped by Patch. Though she was not much more than two-thirds of Whiteblood’s size, the infant clinging to her chest was like a badge of authority. Whiteblood took one fruit and, grumbling, moved back, giving way to Patch.

While this was going on Roamer, like the brothers, knew that she would get no nearer to the fruit until the dominant ones had taken what they wanted.

Alone, she walked carefully, all four limbs grasping, toward the edge of the raft, where the tangle of branches was a little looser. The two terrified crowders, huddled together, skittered away as she approached. Through the foliage she could see murky brown water, littered with bits of wood and leaf, rippling languidly. The sun glimmered in a hundred places, shining through gaps in the cover of the fallen tree, and the dancing light was entrancing, distracting.

Roamer was hungry, but she was also thirsty. She dipped her hand cautiously into the water — it was cool — and scooped up a mouthful. The water was mildly salty — not bitterly so, for even so far from land the river’s powerful outflow diluted the ocean’s brine. But as she drank the taste of salt began to build up in her mouth, and she spat out her last mouthful.

Hungry, bored, the brothers came to inspect her as she drank, head bent down into the foliage, arm outstretched, buttocks raised. They sniffed her curiously, but they could smell how young she was, too young to mate.

When the older ones were done, Roamer and the others fell on the fruit.

With their bellies full for now, the anthros were calming down. But already the haphazard raft had drifted out of sight of the land, already the anthros had eaten much of the fruit from the drowned mango tree. And already the potbelly, complacently munching, had stripped half the branches of their leaves.

And none of them had seen the pale gray triangle that slid silently through the water, not meters away.

The shark circled the crude, disintegrating raft. Alerted by the feeding frenzy as the drowned inhabitants of the riverbank forest were washed out to the waiting mouths of the ocean, the shark had been attracted by the scent of stale blood that leaked from the indricothere carcass. But now it sensed motion on the tangled foliage that floated overhead. It circled, calculating, patient.

The shark was not as intelligent as its parallels on land. But then it was not much like an animal at all. The bones of its back were not bone, but tough cartilage that gave the shark better flexibility than more advanced fish. Its jaw was cartilage too, in which were loosely attached teeth, serrated like steak knives, perfect for shearing flesh. Its projecting snout looked crude, but it cut through the water with the precision of a submarine’s engineering, and it was equipped with nostrils that could detect minute traces of blood. Beneath the snout was a special organ with extraordinary sensitivity to vibration, enabling it to sense the struggles of a frightened animal across immense distances. Behind its small head, the shark’s entire body was made of muscle, designed for power, for forward drive. It was like a battering ram.

Sharks had already been the ocean’s top predators for three hundred million years. They had endured through the great extinctions, while families of land predators had come and gone. They had seen off competition from new classes of animals, some much younger, like the true fish. Over that vast period of time, the sharks’ body design had barely modified, for there was no need.